All consonants are geminated following schwa.
/lˀ ɽ/ are derived diachronically from / ʔl ʔr/. /ⱱ/ is derived from /ʔv/ and /ʔb/. /ɽ/ may still be synchronically analyzable as /ʔr/, but the other two consonants must be taken as phonemic.
Donohue writes /ˀl/ instead of /lˀ/.